Dr. Zheguang Lin is an Associate Professor at Yangzhou University, China. He completed his PhD training at Zhejiang University and served as a visiting scholar at Agroscope. His research focuses on honey bee genetics, breeding, and molecular mechanisms of bee health, with the first proposal of the concept of honey bee breed. He has published dozens of peer-reviewed papers in SCI journals such as Science Advances, PLoS Genetics, and Conservation Letters. He has won international and national academic awards for his research and teaching work.

ORCID: 0000-0003-4270-9360

ResearchGate: https://www.researchgate.net/profile/Zhe-Guang-Lin

Selected Publications  ^#equal contribution  ^*correspondence

Lin Z^#, Zhu Z^#, Zhuang M^#, Wang Z, Zhang Y, Gao F, Niu Q^*, Ji T^*. Effects of local domestication warrant attention in honey bee population genetics.

Sci Adv 2023; 9(18): eade7917. 

Lin Z^*, Zhang Y, Wang K, Jiang X^*, Ji T^*, 2025. Genetic bottlenecks in modern beekeeping: Implications for conservation and sustainable pollination.

Conserv Lett 2025; 18(6): e13156. 

Lin Z^#, Wang S^#, Neumann P, Chen G, Page P, Li L, Hu F, Zheng H^*, Dietemann V. Population genetics and host specificity of Varroa destructor mites infesting Eastern and Western honeybees.

J Pest Sci 2021; 94: 1487-1504. 

Lin Z, Shen S, Wang K, Ji T^*. Biotic and abiotic stresses on honeybee health.

Integr Zool 2023; 19(3): 442-457. 

Lin Z^*, Yang L, Wang Z, Wang K, Niu Q^*, Ji T^*. Honey bee breeding and breed: Advancements, challenges, and prospects.

Anim Res One Heal 2025; 3(4): 350-357.

Li X, Yang X, You F, Miao C, Li M, Wang K, Niu Q, Ji T, Wang Z^*, Lin Z^*. Differences between uncapping and removal behaviors in Apis cerana from the perspective of long non-coding RNAs.

BMC Genomics 2024; 25: 912.

Shen S, Li M, Dong T, Xu K, Wu L, Wang K, Niu D, Niu Q, Ji T, Wang Z, Lin Z^*. Gene expression of long non-coding RNAs and messenger RNAs in Varroa destructor mites exposed to essential oil of camphor leaves.

Exp Appl Acarol 2025; 95: 45.

Dong T, Lv Q, Wu L*, Yang L, Liang Z, Miao C, Zhang Y, Wang K, Niu Q, Ji T, Lin Z*. Morphological and genetic variation of the Chinese honey bee (Apis cerana cerana Fabricius, 1793) in Wanyuan, Southwest China.

Insects 2026; 17(2): 189.

Lin Z, Zheng M, Li Z, Ji T^*. Editorial: Biotic and abiotic stresses on honeybee physiology and colony health.

Front Physiol 2023; 14: 1260547.

Lin Z, Zhang N, Wang Z, Zhuang M, Wang Q, Niu D, Page P, Wang K, Niu Q^*, Ji T^*. Acute and chronic viruses mediated by an ectoparasite target different developmental stages of honeybee (Apis mellifera and Apis cerana) brood.

Front Vet Sci 2022; 9: 951159.

Lin Z, Xu H, Su X, Ke Y, Wang W, Li Y, Zhuang M, Chen H, Liu Y, Wang K, Chen G, Ji T^*. Investigation of circular RNAs in an ectoparasitic mite Varroa destructor (Acarina: Varroidae) of the honey bee.

Parasitol Res 2021; 120: 715-723.

Lin Z, Zhao L, Cao L, Xu H, Miao C, Jie H, Zhang Z, Liu Y, Gao F, Wang Z, Chen G, Ji T^*. A survey of common pathogens of Apis spp. in wild non-cave honeybees in Southwest China.

J Wildl Dis 2021; 57(4): 932-935.

Lin Z, Chen H, Page P, Wang K, Ji T, Chen G^*. First record of pseudoscorpions in the Eastern honey bee colonies in China.

J Apic Res 2020; 59(4): 515-518.

Lin Z, Liu Y, Chen X, Han C, Wang W, Ke Y, Su X, Li Y, Chen H, Xu H, Chen G, Ji T^*. Genome-wide identification of long non-coding RNAs in the gravid ectoparasite Varroa destructor.

Front Genet 2020; 11: 575680.

Lin Z, Su X, Wang S, Ji T, Hu F, Zheng H^*. Fumigant toxicity of eleven Chinese herbal essential oils against an ectoparasitic mite (Varroa destructor) of honey bee (Apis mellifera).

J Apic Res 2020; 59(2): 204-210.

Lin Z^#, Qin Y^#, Page P, Wang S, Li L, Wen Z, Hu F, Neumann P, Zheng H^*, Dietemann V. Reproduction of parasitic mites Varroa destructor in original and new honey bee hosts.

Ecol Evol 2018; 8: 2135-2145.

Zhu K^#, Liu M^#, Fu Z^#, Liang H, Zhou Z, Kong Y, Lin Z, Luo J, Zheng H, Wan P, Zhou Z, Zhang J, Zen K, Chen J^*, Hu F^*, Zhang C^*, Ren J^*, Chen X^*. Plant microRNAs in larval food regulate honey bee caste development.

PLoS Genet 2017; 13(8): e1006946.

Lin Z^#, Page P^#, Li L, Qin Y, Zhang Y, Hu F, Neumann P, Zheng H^*, Dietemann V. Go East for better honey bee health: Apis cerana is faster at hygienic behavior than A. mellifera.

PLoS One 2016; 11(9): e0162647.

Page P^#, Lin Z^#, Buawangpong N^#, Zheng H^*, Hu F, Neumann P, Chantawannakul P^*, Dietemann V. Social apoptosis in honey bee superorganisms.

Sci Rep 2016; 6: 27210.

Research Project Leader

National Natural Science Foundation of China (NSFC) General Program, No. 32272935, January 2023–December 2026

National Natural Science Foundation of China (NSFC) Youth Program, No. 31902220, January 2020–December 2022

Excellent Youth Foundation of Jiangsu Province, No. BK20240158, September 2024–August 2027

Young Elite Scientists Sponsorship Program by Jiangsu Association, No. TJ-2023-095, July 2023–June 2025

Entrusted Project of the Ministry of Agriculture and Rural Affairs of China, November 2021–November 2022

China Postdoctoral Science Foundation General Program, No. 2019M651983, January 2019–December 2020

Qinglan Project for Mid-Career Academic Leaders of Yangzhou University, July 2025–July 2028

Qinglan Project for Outstanding Young Faculty of Yangzhou University, July 2024–July 2027

Teaching Reform Research Project of Yangzhou University (Special Project for Innovation and Entrepreneurship Education Reform), No. yzucx2023-2B, December 2023–November 2025

Cross-Disciplinary High-Level Young Talent Cultivation Project of Yangzhou University, November 2022–October 2023

Lvyangjinfeng Program of Yangzhou, No. YZLYJFJH2021YXBS158, January 2022–December 2024

Teaching Reform Research Project of Yangzhou University (Doctoral Special Project), No. YZUJX2022—C15, September 2022–August 2024

High-Level Talent Research Start-Up Project of Yangzhou University, April 2021–March 2027

College Student Scientific and Technological Innovation Project of Zhejiang Province (Xinmiao Talent Program), No. 2015R401266, June 2015–May 2016